Grant Information

CHARACTERIZATION OF A PUTATIVE INSECT-TRANSMISSION DETERMINANT/VIRULENCE GENE (HYPL) OF ¿CANDIDATUS LIBERIBACTER ASIATICUS¿

Sponsoring Institution Agricultural Research Service/USDA
Status ACTIVE
Funding Source USDA INHOUSE
Reporting Frequency Annual
Project Director DUAN Y
Accession Number 419962
Project Number 6034-22000-039-04R
Dates 2010-05-01 - 2013-03-31
Recipient Organization AGRICULTURAL RESEARCH SERVICE
219 SOUTH ROCK ROAD
FT PIERCE,FL 34945
Keywords asiaticus
candidatus
citrus
gene
huanglongbing
hypl
liberibacter
Research Effort Applied (30%)
Basic (60%)
Developmental (10%)
Classification Parameters
Knowledge AreaSubject of InvestigationField of SciencePercent
212 - Pathogens and Nematodes Affecting Plants 910 - Grapefruit 1040 - Molecular biology 50%
212 - Pathogens and Nematodes Affecting Plants 920 - Orange 1060 - Biology (whole systems) 50%
Goals / Objectives

Characterize the hypl gene and determine its effects on insect transmission and/or virulence in host plants. The objectives of the original proposal include:

  1. determine the cellular localization of the Hypl protein within insect or plant cells and the efficacy of the two putative NLSs coded by the hypl gene
  2. determine the expression dynamics of hypl both in psyllids and plants (citrus and periwinkle) and elucidate its activity with respect to effector function
  3. develop new techniques for more sensitive detection and differentiation of Las bacteria based on the unique sequence and antigenicity of the hypl gene
  4. develop transgenic citrus resistant to HLB by expressing plantibody against the Hypl protein
Methods (unparsed)

Transient expression with alternative expression systems and RT-qPCR, etc., will be used to elucidate the function of the hypl gene of Las and shed the light on the molecular mechanism of this "phase variation" phenomenon and thereby develop a novel control strategy for citrus HLB. In addition, antibodies and probes along with standardized protocols, developed during this project can be applied for better detection and differentiation of the HLB bacteria.

Methods
Transient expression with alternative expression systems and RT-qPCR, etc., will be used to elucidate the function of the hypl gene of Las and shed the light on the molecular mechanism of this "phase variation" phenomenon and thereby develop a novel control strategy for citrus HLB. In addition, antibodies and probes along with standardized protocols, developed during this project can be applied for better detection and differentiation of the HLB bacteria.
Project Timeline Tracking

Outputs

Transient expression with alternative expression systems and RT-qPCR, etc. , will be used to elucidate the function of the hypl gene of Las and shed the light on the molecular mechanism of this "phase variation" phenomenon and thereby develop a novel control strategy for citrus HLB. In addition, antibodies and probes along with standardized protocols, developed during this project can be applied for better detection and differentiation of the HLB bacteria. This research relates to inhouse project objective: 1. Characterize ecology, biology, epidemiology, molecular genetics, and vector and host (crop and weed) interactions of domestic, exotic, newly emerging, and re- emerging pathogens, and 2. Develop/refine rapid, sensitive reliable detection/sampling methods for pathogens. We have characterized two novel autotransporter proteins of �Candidatus Liberibacter asiaticus� (Las), and redesignated them as LasAI and LasAII in lieu of the previous names HyvI and HyvII. Bioinformatic analyses revealed that LasAI and LasAII share the structural features of an autotransporter family containing large repeats of a passenger domain and a unique C-terminal translocator domain. When fused to the Green fluorescent protein (GFP) gene and expressed in Escherichia coli, the LasAI C-terminus and the full length LasAII were localized to the bacterial poles, similar to other members of autotransporter family. Despite the absence of the signal peptide, LasAI was found to localize at the cell surface by immuno-dot blot using a monoclonal antibody against the partial LasAI protein. Its surface localization was also confirmed by the removal of the LasAI antigen using a proteinase K treatment of the intact bacterial cells. When co-inoculated with a P19 gene silencing suppressor and transiently expressed in tobacco leaves, the GFP-LasAI translocator targeted to the mitochondria of host plants. This is the first report that Las encodes novel autotransporters that target to mitochondria.